What is the role of the destination directory's ownership in causing a 500 error?

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Grade: Education Subject: Support
What is the role of the destination directory's ownership in causing a 500 error?
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The destination directory must be owned by the web server user (e.g., 'apache' or 'www-data'). If the directory is owned by a different user, the web server won't have the necessary permissions to write to it, resulting in a 500 error.